Olmstead et al. (1999) presented phylogenetic analyses of the Solanaceae based on restriction site variation of the entire chloroplast genome (Olmstead & Palmer, 1992) and DNA sequences of two genes, rbcL and ndhF (Olmstead et al. 1992, 1994; Olmstead & Sweere, 1994). These analyses contained one species of Nolana, and Nolana was deeply nested within the Solanaceae-Solanoideae in a clade with Lycium and Grabowskia (Lycieae). Johnston (1936), on morphological evidence, already had suggested that Nolana shared relationships with both Lycium and Grabowskia.

Tortuga Bay - Isla Santa Cruz


Grabowskia boerhaviifolia is an autodisjunction from
the mainland of northern Peru.

Grabowskia boerhaviifolia usually inhabits arid and near shore environments.
and its leaves are laminar, elliptic-ovate and succulent.

Lycium minimum is a Galápagos Island endemic growing here on the west
side of Santa Curz's Tortuga Bay.

Lycium minimum with only four petals and anthers

Lycium minimum on Isla Floreana / Santa María

Lycium minimum growing directly from lava on Floreana (20 February 2004)

Lycium minimum along the beach of Punta Cormorant (20 February 2004).

Lycium minimum is found in near ocean sites growing in
either sand or directly from lava

Lycium minimum has highly succulent leaves and thorny twigs.
It was not flowering at this date (20 Feb 2004).

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